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After months of peace, the cities of Tollen and Montevado suddenly and mysteriously disappear underground, leading the Coalition of Ordered Governments (COG) to suspect the resurgence of the Locust. Soon after, the once impenetrable Jacinto, one of the last remaining safe havens for humans, begins to show signs that the same fate awaits it. Though most of the underground army of the Locust Horde was destroyed, the explosion also caused much of the liquid Imulsion underground to vaporize, causing a fatal disease called Rustlung to spread among the diminished human population.

With the COG army disbanded, the remnant must band together to survive against a new threat, the Lambent.Gears of War 2 takes place six months after the detonation of the Lightmass Bomb at the end of the first game. Gears of War 3 takes place 2 years after the sinking of Jacinto in Gears 2. The sequel Gears of War 2 takes place six months after the events of the first game, and follows Delta Squad and the COG army as it launches an assault directly against the Hollow in an effort to end the war. In Gears of War, the protagonist Marcus Fenix, a soldier or "Gear" in the Coalition of Ordered Governments, leads his squad on a mission to deploy the Lightmass bomb and destroy the Locust Horde in their underground strongholds.

All of the games are noted for their heavy violence. The three games that currently make up the series take place on the fictional planet Sera and focus on a war between humans and creatures known as Locust. Gears of War is a third-person shooter video game franchise created and owned by Epic Games and published by Microsoft Studios. Like all members of his family, he viewed dying for the COG as the highest honor.

Anthony, unlike the veteran Gears, preferred full armor, which some claim led to his death. Though he had great respect for veteran Gears like Marcus Fenix, he held some ignorance towards the carnage of the Pendulum Wars and the battle of Aspho Fields, viewing it as "cool". He was an enthusiastic soldier with a gung-ho attitude willing to rush into enemy fire.